- Leaves simple, opposite, serrate or crenate.
- Sheath-like interpetiolar stipule with fine teeth.
- Aromatic odor.
- Flowers unisexual, reduced and primitive, males on axillary catkins, females apetalous.
- Fruit a small, white, and berry-like.
Diagnosis: Small trees or shrubs, aromatic odor, opposite serrate leaves, interpetiolar sheath-like stipule with fine teeth.
Local distribution: Cloud forest ridges and Atlantic slope rain forest.
|Gentianaceae||: margin entire, no teeth on interpetiolar sheath.
|Rubiaceae||: margin entire, without aromatic odor.
Diversity: Total: 1 genus, 4 species. Trees: 1 genus, 3 species.
Comments: Considered to be one of the most primitive angiosperm families.